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Ecology

Guidelines for the Selection of Biological Heritage Sites

Pied Flycatcher
Contents

7. Section 2: Species Guidelines

7.8 INVERTEBRATES: BUTTERFLIES AND MOTHS (LEPIDOPTERA)

Application (all invertebrate guidelines)

Identification of sites on the basis of their invertebrate animal species should take account of the needs of many invertebrates for habitat and structural diversity, both at the 'macro' and 'micro' scales.  Small-scale habitats of particular importance to some invertebrates, such as dead wood or small patches of bare ground, occur and indeed may shift their position over time within large scale habitats e.g. woodland or grassland, on which the invertebrates also depend.  Moreover the larval and adult phases of the same species often need quite different habitats.  Invertebrates generally have annual life cycles, and their survival on a site depends on the continued availability of the right mixture of habitats at the right time of year - every year.  Such factors should be taken into account when determining site boundaries.

Species lists, especially for less well-recorded groups of invertebrates, should be regarded as tentative.  They include species recorded in Lancashire since 1950, those not recorded since 1978 being shown with an asterisk, or other notation.

It should be noted that the category 'nationally scarce', referring to species believed to occur between 16 and 100 10km squares of the National Grid, is sometimes subdivided in relation to invertebrate groups into 'Notable A' and 'Notable B'.  This subdivision has not been recognised in the following guidelines.

Justification

In Lancashire, as elsewhere, there are more species of invertebrate animals than of all plants and other animals combined.  Many invertebrate groups and species have declined dramatically in recent decades, and their conservation is a matter of widespread concern in Europe.  Until recently, it was thought that if sites were selected (and managed) on the basis of their botanical interest, then the invertebrates would automatically be catered for too.  This is not so:  it is important that due regard is paid to the contribution that invertebrates make to biodiversity, and of their habitat needs, in their own right, insofar as available information allows.

With such a large number of invertebrate groups, and the relative lack of knowledge about many of these, it is impossible to develop individual sets of guidelines for every species-group at the present time.  Where there is an adequate database for a particular species-group relating to Lancashire, specific guidelines have been devised and appear below.  For the remainder, there is a single, 'catch all', section which follows the standard approach adopted throughout the species section.

i)  BUTTERFLIES AND MOTHS (LEPIDOPTERA)

Application (all butterfly and moth guidelines)

For the purposes of these guidelines acceptable evidence of breeding by butterfly and moth species includes the presence of eggs, or larvae, or pupae, or repeated sightings of adults in suitable habitats.  It should be noted that the lists of moth species are incomplete and, for certain guidelines, not yet available.

Justification (all butterfly and moth guidelines)

Butterflies are popular and conspicuous insects, and are relatively well-recorded.  Moths are much more numerous in terms of species, and less well-recorded.  Each species requires not only the right foodplant and habitat for its larvae but also suitable habitat and nectar-producing flowers for the adults.  More than 20% of all British butterfly species are regarded as threatened in a national context.

Le1 Any site which regularly supports a breeding population of a species of butterfly or moth included in British Red Data Books: 2.  Insects. (1)

Application

The species to which this guideline applies include:


BUTTERFLIES
Argynnis adippe
High Brown Fritillary
MOTHS
* Homoeosoma nimbella
Small clouded Knot-horn
Lycia zonaria
Belted Beauty
Photedes captiuncula
Least Minor

High Brown Fritillary

Justification

The species in the above category are threatened or rare in Britain and there is a national responsibility for their conservation.  The high brown fritillary, in particular, has suffered a dramatic decline in Britain since 1950.  There is an urgent national need to conserve its few remaining populations, some of the most important of which are in Lancashire.


Le2 Any site which regularly supports a breeding population of a 'nationally scarce' species of butterfly or moth.

Application

The species to which this guideline applies include:


BUTTERFLIES
Aricia agestis
Brown Argus
Aricia artaxerxes
Northern Brown Argus
Boloria euphrosyne
Pearl-bordered Fritillary
Coenonympha tullia
Large Heath
* Erebia aethiops
Scotch Argus
Hamearis lucina
Duke of Burgundy
MOTHS
Actebia praecox
Portland Moth
Adscita geryon
Cistus Forester
* Agrotis ripae
Sand Dart
Anania funebris
White-spotted Sable
Atolmis rubricollis
Red-necked Footman
* Carsia sororiata
Manchester Treble-bar
* Catarhoe rubidata
Ruddy Carpet
Chilodes maritimus
Silky Wainscot
Chlorissa viridata
Small Grass Emerald
Crambus ericella
Heath Grass-veneer
* Crambus hamella
Pearl-streak Grass-veneer
Deltolte uncula
Silver Hook
Discoloxia blomeri
Blomer's Rivulet
Eana penziana
A tortrix moth
* Eriogaster lanestris
Small Eggar
Eucosmomorpha albersana
A tortrix moth
* Eudonia delunella
Resin Grey
Eudonia lineola
A pyralid moth
Eupithecia expallidata
Bleached Pug
Eurrhypara terrealis
Northern Pearl
* Euxoa cursoria
Coast Dart
* Hypenodes humidialis
Marsh Oblique-barred
Idaea muricata
Purple-bordered Gold
* Microstega pandalis
Bordered Pearl
* Mythimna litoralis
Shore Wainscot
* Olethreutes olivana
A tortrix moth
* Pechipogon strigilata
Common Fan-foot
* Pediasia aridella
Saltmarsh Grass-veneer
Perconia strigillaria
Grass Wave
Perizoma minorata ssp. ericetata
Heath Rivulet
Perizoma taeniata
Barred Carpet
Phalonidia curvistrigana
A micro-moth
* Rheumaptera hastata
Argent and Sable
* Scoparia ancipitella
Elm Grey
Sideridis albicolon
White Colon
Sitochroa palealis
A pyralid moth
Thera juniperata
Juniper Carpet
Tetheella fluctuosa
Satin Lutestring
Trichopteryx polycommata
Barred Tooth-striped
* Xylena exsoleta
Sword Grass

Justification

Nationally scarce species are recorded only from 16-100 10km squares (inclusive) in Britain; there is a national responsibility to ensure their conservation.


Le3 Any site which regularly supports a breeding population of a species of butterfly or moth which occurs at 3 or fewer localities in Lancashire.

Application

All sites for species in the above category which are not included in Guidelines Le1 or Le2 should be considered.

The species to which this guideline applies include:


BUTTERFLIES
Argynnis aglaja
Dark Green Fritillary
MOTHS
List of species not available.

Justification

Although more widespread nationally than those species listed under Guidelines Le1 and Le2, this species is very rare in Lancashire.


Le4 Any site which regularly supports a breeding population of a species of butterfly or moth which is recorded from more than three localities in Lancashire, but which could be at risk because of small populations, habitat loss or change, or is at the edge of its British range, where such populations contribute significantly to the distribution pattern or the total population size of that species in the County.

Application

Sites for butterflies and moths in the above categories (not included under Guidelines Le1 or Le2) may be considered for inclusion where they significantly extend the geographical range of the species in Lancashire, or support a significant proportion of the estimated total County population of that species.  The species to which the guideline applies include:


BUTTERFLIES
Boloria selene 
Small Pearl-bordered Fritillary
Callophyrys rubi
Green Hairstreak
Erynnis tages
Dingy Skipper
Gonepteryx rhamni
Brimstone
Hipparchia semele
Grayling
Pararge aegeria 
Speckled Wood
Polygonia c-album
Comma
Quercusia quercus
Purple Hairstreak
Strymonidia w-album
White-letter Hairstreak
MOTHS
List of species not available.

Justification

Species included here, whilst not as rare in Lancashire as those under Guidelines Le1, Le2 or Le3, are nevertheless either of very restricted distribution or exist only as small scattered populations in the County.


Le5 Any site which regularly supports breeding populations of 9 or more butterfly species (excluding those species which are migratory or are largely associated with cultivated plants).

Brimstone

Application

The following species are relevant for the purposes of this guideline:


BUTTERFLIES
Aglais urticae
Small Tortoiseshell
Anthocharis cardamines
Orange Tip
Boloria selene
Small Pearl-bordered Fritillary
Callophrys rubi
Green Hairstreak
Celastrina argiolus
Holly Blue
Coenonympha pamphilus
Small Heath
Erynnis tages
Dingy Skipper
Gonepteryx rhamni
Brimstone
Hipparchia semele                
Grayling
Inachis io
Peacock
Lasiommata megera              
Wall
Lycaena phlaeas
Small Copper
Maniola jurtina
Meadow Brown
Ochlodes venata
Large Skipper
Pararge aegeria
Speckled Wood
Pieris napi
Green-veined White
Polyommatus icarus
Common Blue
Pyroia tithonus
Gatekeeper
Quercusia quercus                
Purple Hairstreak
Strymonidia w-album
White-letter Hairstreak
Thymelicus sylvestris
Small Skipper
MOTHS
List of species not available

Justification

The purpose of this guideline is to identify sites with significant habitat and structural diversity, not involving crop or cultivated plants, which support  notable assemblages of breeding butterflies.


(1) Shirt (1987)
* Last recorded 1950-1977

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